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It also contains a sulfur molecule, which plays a major role in its antioxidant and detoxification functions

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doi:10.1016/S0753-3322(03)00043-X Ubiyvovk VM, Ananin VM, Malyshev AY, Kang HA, Sibirny AA (2011) Optimization of glutathione production in batch and fed-batch cultures by the wild-type and recombinant strains of the methylotrophic yeast Hansenula polymorpha DL-1